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1977-1983: Medical studies at the universities of Essen, Tufts (Boston) and Göttingen.
1983-1984: Clinical electives ("Praktisches Jahr") in Adelaide, Boston and Montreal.
June 1984: Medical dissertation ("Dr. med.") at the Max Planck Institute for Biophysical Chemistry, Dept. of Neurobiology (supervisor: O. Creutzfeldt). Visual Psychophysics and Neurophysiology.
1985-1988: Graduate Studies ("Ph.D.") at the Montreal Neurological Institute, McGill University, Montreal/Canada (supervisors: L. Yamamoto and M. Diksic). Deoxyglucose mapping of sensory cortex.
1988 - 1990: Post-Doc in the Dept. of Brain and Cognitive Sciences (with Ron McKay), M.I.T., Cambridge/USA. Immortalization of neural precursor cells.
1990 - 1991: Post-Doc in the Dept. of Biophysics (with Masatoshi Takeichi), Kyoto University, Kyoto/Japan. Cadherins and brain development.
1991 - 1995: Post-Doc in the Dept. of Biochemistry (head: Uli Schwarz), Max Planck Institute for Developmental Biology, Tübingen/Germany.
1995 - 1997: Group Leader at the Institute of Biology III, University of Freiburg, Freiburg/Germany.
1997 - 2003: Associate Professor (C3) of Anatomy, University of Essen Medical School, Essen/Germany.
2003-2021: Professor of Anatomy (C4/W3) and Head, Institute of Anatomy I, University of Jena School of Medicine, Jena/Germany
2008: Declined an offer for a Professor of Anatomy (W3) position at the University of Düsseldorf.
March-July 2015: Visiting Fellow, Laboratory of Experimental Psychology, University of Leuven (KUL), Leuven/Belgium (with J. Wagemans)
2018-2024: Curator, Anatomical Collection, University of Jena School of Medicine
since 2021: Emeritus Professor, University of Jena School of Medicine
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